CAS Ichthyology Primary Types Imagebase

SU 6480 Aemasia lichenosa Jordan [D. S.] & Snyder [J. O.], 1901 Holotype  [Collection database record] No indication that fig. is based on type.
lichenosa, Aemasia Jordan [D. S.] & Snyder [J. O.] 1901:883, Fig. 20 [Proceedings of the United States National Museum v. 23 (no. 1239); ref. 2508] Wakanoura, Wakayama Prefecture, Japan, Inland Sea, Western North Pacific. Holotype: CAS-SU 6480. Paratypes: USNM 49976 (1). Type catalog: Böhlke 1953:46 [ref. 12291], Smith 1994:14 [ref. 21917], Böhlke & Smith 2002:121 [ref. 26380]. Types: Shinohara & Williams 2006:548 [ref. 31656]. Galápagos population may be a separate species -- Smith 2012:8 [ref. 32155]. •Valid as Enchelycore lichenosa (Jordan & Snyder 1901) -- (Hatooka in Masuda et al. 1984:23 [ref. 6441], Böhlke et al. 1989:135 [ref. 13286], Allen & Robertson 1994:44 [ref. 22193], Chen et al. 1994:47 [ref. 22258], Grove & Lavenberg 1997:159 [ref. 24023], Nakabo 2000:200 [ref. 25086], Böhlke & McCosker in Randall & Lim 2000:585 [ref. 25122], Nakabo 2002:200 [ref. 26001], Böhlke & Smith 2002:121 [ref. 26380], Smith 2002:4 [ref. 26440], Shibukawa et al. 2007:187 [ref. 29475], McCosker & Rosenblatt 2010:188 [ref. 30957], Zhang et al. 2010:252 [ref. 31511], Smith 2012:8 [ref. 32155], Ho et al. 2015:153 [ref. 34130], Mohapatra et al. 2017:598 [ref. 35581], Ramos-Castro et al. 2020:83 [ref. 37651], Dale et al. 2021:312 [ref. 38298], Oomori et al. 2022:26 [ref. 39429]). Current status: Valid as Enchelycore lichenosa (Jordan & Snyder 1901). Muraenidae: Muraeninae. Distribution: Northwestern Pacific: Japan; eastern Pacific: Panama south to Colombia, including Galápagos Islands (Ecuador). Habitat: marine.
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© 2001 California Academy of Sciences

SU 6481 Echidna kishinouyei Jordan [D. S.] & Snyder [J. O.], 1901 Holotype  [Collection database record]
kishinouyei, Echidna Jordan [D. S.] & Snyder [J. O.] 1901:885, Fig. 21 [Proceedings of the United States National Museum v. 23 (no. 1239); ref. 2508] Okinawa Island, Ryukyu Islands, Japan. Holotype (unique): CAS-SU 6481. Type catalog: Böhlke 1953:46 [ref. 12291], Böhlke & Smith 2002:119 [ref. 26380]. •Synonym of Gymnothorax thyrsoideus (Richardson 1845) -- (Böhlke & Smith 2002:119 [ref. 26380], Smith 2012:28 [ref. 32155]). Current status: Synonym of Gymnothorax thyrsoideus (Richardson 1845). Muraenidae: Muraeninae. Habitat: marine.
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© 2001 California Academy of Sciences

CAS 35250 Gymnothorax breedeni McCosker [J. E.] & Randall [J. E.], 1977 Holotype  [Collection database record]
breedeni, Gymnothorax McCosker [J. E.] & Randall [J. E.] 1977:162, Figs. 1-3 [Proceedings of the California Academy of Sciences (Series 4) v. 41 (no. 3); ref. 7267] Itsandra, Grande Comore Island, Comoro Islands, Indian Ocean. Holotype: CAS 35250. Paratypes: ANSP 134222 (1), BPBM 11903 (1), USNM 215283 (1). Type catalog: Böhlke 1984:130 [ref. 13621], Smith 1994:17 [ref. 21917], Böhlke & Smith 2002:98 [ref. 26380]. •Valid as Gymnothorax breedeni McCosker & Randall 1977 -- (Castle & McCosker 1986:168 [ref. 5645], Myers 1999:46 [ref. 23965], Böhlke et al. 1999:1656 [ref. 24692], Allen 2000:84 [ref. 37268], Randall & Earle 2000:6 [ref. 25806], Böhlke & Smith 2002:98 [ref. 26380], Randall 2005:39 [ref. 28239], Randall et al. 2005:118 [ref. 28745], Allen & Erdmann 2012:81 [ref. 31980], Smith 2012:12 [ref. 32155], Fricke et al. 2013:251 [ref. 32706], Delrieu-Trottin et al. 2015:4 [ref. 33988], Koeda & Akita 2017:32 [ref. 35353], Eudeline 2022:136 [ref. 39191], Smith & Böhlke 2022:48 [ref. 39681], Allen & Erdmann 2024:90 [ref. 40796]). Current status: Valid as Gymnothorax breedeni McCosker & Randall 1977. Muraenidae: Muraeninae. Distribution: Indo-West Pacific: KwaZulu-Natal (South Africa), East Africa (Mozambique), Seychelles, Comoros and Mayotte (France) east to Caroline Islands and Marquesas Islands (French Polynesia), north to southern Japan. Habitat: marine.
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© 2001 California Academy of Sciences

CAS 48815 Gymnothorax enigmaticus McCosker [J. E.] & Randall [J. E.], 1982 Holotype  [Collection database record]
enigmaticus, Gymnothorax McCosker [J. E.] & Randall [J. E.] 1982:18, Figs. 1, 4 [Proceedings of the California Academy of Sciences (Series 4) v. 43 (no. 2); ref. 8445] Off Ngatchab Beach, Angaur Island, Palau Islands, Western Caroline Islands, western Pacific. Holotype: CAS 48815. Paratypes: BPBM 7715 (2), 8184 (1), 9491 (1), 12354 (1), 22339 (2), 22827 (1), 22809 (2); CAS 42377 (1), 48816-26 (1, 1, 1, 1, 1, 3, 2, 8, 2, 1, 1). Type catalog: Böhlke & Smith 2002:108 [ref. 26380] with holotype as 4881. •Valid as Gymnothorax enigmaticus McCosker & Randall 1982 -- (Hatooka in Masuda et al. 1984:23 [ref. 6441], Castle & McCosker 1986:169 [ref. 5645], Winterbottom et al. 1989:8 [ref. 13251], Quéro & Saldanha 1995:66 [ref. 22017], Allen 1997:54 [ref. 23977], Myers 1999:46 [ref. 23965], Fricke 1999:45 [ref. 24106], Böhlke et al. 1999:1656 [ref. 24692], Allen 2000:84 [ref. 37268], Compagno in Randall & Lim 2000:585 [ref. 25122], Nakabo 2000:205 [ref. 25086], Böhlke & McCosker 2001:73 [ref. 25226], Nakabo 2002:205 [ref. 26001], Böhlke & Smith 2002:108 [ref. 26380], Allen & Adrim 2003:23 [ref. 26830], Manilo & Bogorodsky 2003:S95 [ref. 27377], Myers & Donaldson 2003:611 [ref. 27495], Lobel & Lobel 2004:68 [ref. 27576], Randall et al. 2004:6 [ref. 27624], Randall 2005:40 [ref. 28239], Allen et al. 2006:249 [ref. 29005], Randall 2007:92 [ref. 30952], Fricke et al. 2009:15 [ref. 30213], Motomura et al. 2010:71 [ref. 31256], Allen & Erdmann 2012:83 [ref. 31980], Smith 2012:14 [ref. 32155], Fricke et al. 2014:18 [ref. 33932], Fricke et al. 2018:35 [ref. 35805], Ho et al. 2018:12 [ref. 36050], Arndt & Fricke 2019:23 [ref. 36883], Smith & Böhlke 2022:50 [ref. 39681], Allen & Erdmann 2024:92 [ref. 40796]). Current status: Valid as Gymnothorax enigmaticus McCosker & Randall 1982. Muraenidae: Muraeninae. Distribution: Indo-West Pacific: Gulf of Aden, East Africa, KwaZulu-Natal (South Africa), Aldabra (Seychelles), Madagascar and western Mascarenes (La Réunion, Mauritius) east to Tuamotu Archipelago (French Polynesia), north to Taiwan and southern Japan, south to Timor Sea and Queensland (Australia), and Samoa. Habitat: marine.
Not Illustrated
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© 2001 California Academy of Sciences

SU 12686 Gymnothorax pescadoris Jordan [D. S.] & Evermann [B. W.], 1902 Holotype  [Collection database record]
pescadoris, Gymnothorax Jordan [D. S.] & Evermann [B. W.] 1902:326, Fig. 8 [Proceedings of the United States National Museum v. 25 (no. 1289); ref. 2447] Kotosho, Hokoto[Hokuto, Taipai City] or Pescadores Islands, Taiwan. Holotype (unique): CAS-SU 12686 [ex CAS-SU 7131]. Type catalog: Böhlke 1953:47 [ref. 12291], Böhlke & Smith 2002:136 [ref. 26380], Ho & Shao 2011:21 [ref. 31432]. •Valid as Gymnothorax pescadoris Jordan & Evermann 1902 -- (Hatooka in Masuda et al. 1984:24 [ref. 6441]). •Synonym of Gymnothorax favagineus Bloch & Schneider 1801 -- (Chen et al. 1994:46, 50 [ref. 22258], Zhang et al. 2010:286 [ref. 31511]). •Synonym of Gymnothorax isingteena (Richardson 1845) -- (Böhlke & Smith 2002:136 [ref. 26380], Ho & Shao 2011:21 [ref. 31432], Smith 2012:17 [ref. 32155], Ho et al. 2015:7 [ref. 34116], Ho et al. 2015:157 [ref. 34130]). Current status: Synonym of Gymnothorax isingteena (Richardson 1845). Muraenidae: Muraeninae. Habitat: marine.
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© 2001 California Academy of Sciences

SU 9215 Gymnothorax philippinus Jordan [D. S.] & Seale [A.], 1907 Holotype  [Collection database record]
philippinus, Gymnothorax Jordan [D. S.] & Seale [A.] 1907:7, Fig. 2 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 606) (for 1906); ref. 2498] Cavite, Luzon Island, Philippines. Holotype (unique): CAS-SU 9215. Type catalog: Böhlke 1953:47 [ref. 12291], Böhlke & Smith 2002:137 [ref. 26380]. See Böhlke et al. 1999:1657 [ref. 24692] for remarks. •Valid as Gymnothorax philippinus Jordan & Seale 1907 -- (Hatooka in Masuda et al. 1984:24 [ref. 6441], Compagno in Randall & Lim 2000:585 [ref. 25122], Böhlke & Smith 2002:137 [ref. 26380] with some doubt, Allen & Erdmann 2012:90 [ref. 31980], Smith 2012:23 [ref. 32155], Allen & Erdmann 2024:100 [ref. 40796]). Current status: Valid as Gymnothorax philippinus Jordan & Seale 1907. Muraenidae: Muraeninae. Distribution: Western Pacific: Philippines, Okinawa and Ryukyu Islands (Japan). Habitat: marine.
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© 2001 California Academy of Sciences

SU 6807 Muraena clepsydra Gilbert [C. H.], 1898 Holotype  [Collection database record]
clepsydra, Muraena Gilbert [C. H.] in Jordan & Evermann 1898:2805 [Bulletin of the United States National Museum No. 47; ref. 2445] (Pacific) Panama. Holotype: CAS-SU 6807. Paratypes: BMNH 1903.5.15.443 (1), CAS-SU 12899-900 (1, 1), USNM 50423 (1). Type catalog: Böhlke 1953:47 [ref. 12291], Smith 1994:21 [ref. 21917], Böhlke & Smith 2002:103 [ref. 26380] Author is C. H. Gilbert. Holotype figured in Gilbert & Starks 1904, Pl. 7 (fig. 13) [ref. 1639]. •Valid as Muraena clepsydra Gilbert 1898 -- (Allen & Robertson 1994:47 [ref. 22193], McCosker & Rosenblatt in Fischer et al. 1995:1312 [ref. 22830], Grove & Lavenberg 1997:158 [ref. 24023], Chirichigno F. & Vélez D. 1998:127 [ref. 24555], Thomson et al. 2000:41 [ref. 25640], Böhlke & Smith 2002:103 [ref. 26380], Nelson et al. 2004:61 [ref. 27807], Love et al. 2005:18 [ref. 37547], McCosker & Rosenblatt 2010:188 [ref. 30957], Smith 2012:31 [ref. 32155], Page et al. 2013:61 [ref. 32708], Fourriére et al. 2016:441 [ref. 34668], Del Moral-Flores et al. 2016:606 [ref. 34923], Love et al. 2021:34 [ref. 39279], Page et al. 2023:52 [ref. 40505]). Current status: Valid as Muraena clepsydra Gilbert 1898. Muraenidae: Muraeninae. Distribution: Eastern Pacific: Pacific coast of Baja California Sur (Mexico) south to northern Peru, including southern Gulf of California (Mexico) and Galápagos Islands (Ecuador). Habitat: marine.
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© 2001 California Academy of Sciences

CAS 4964 Rabula rotchii Clark [H. W.], 1936 Holotype  [Collection database record]
rotchii, Rabula Clark [H. W.] 1936:384 [Proceedings of the California Academy of Sciences (Series 4) v. 21 (no. 29); ref. 839] Tidepools, Black Beach Anchorage, Santa María Island [Charles Island], Galápagos Islands. Holotype (unique): CAS 4964. •Synonym of Echidna nocturna (Cope 1872) -- (McCosker & Rosenblatt 1975:424 [ref. 9341], Smith & Böhlke 2006:36 [ref. 28678], Smith 2012:6 [ref. 32155]). Current status: Synonym of Echidna nocturna (Cope 1872). Muraenidae: Muraeninae. Habitat: marine.
Not Illustrated
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© 2001 California Academy of Sciences

CAS 5744 Uropterygius galapagensis Seale [A.], 1940 Holotype  [Collection database record] Assumed that illustration is based on the type, as only one specimen mentioned in orig. descr.
galapagensis, Uropterygius Seale [A.] 1940:6, Pl. 1 [Allan Hancock Pacific Expedition 1932-40, Los Angeles v. 9 (no. 1); ref. 12332] Post Office Bay, Santa María Island [Charles Island], Galápagos Islands. Holotype (unique): CAS 5744. Type catalog: Böhlke & Smith 2002:159 [ref. 26380]. •Valid as Anarchias galapagensis (Seale 1940) -- (Böhlke et al. 1989:117 [ref. 13286], Allen & Robertson 1994:43 [ref. 22193], McCosker & Rosenblatt in Fischer et al. 1995:1307 [ref. 22830], Grove & Lavenberg 1997:159 [ref. 24023], Thomson et al. 2000:47 [ref. 25640], Böhlke & Smith 2002:159 [ref. 26380], Nelson et al. 2004:60 [ref. 27807], Love et al. 2005:17 [ref. 37547], McCosker & Rosenblatt 2010:188 [ref. 30957], Reece et al. 2010:421 [ref. 30969], Smith 2012:34 [ref. 32155], Page et al. 2013:60 [ref. 32708], Fourriére et al. 2016:441 [ref. 34668], Del Moral-Flores et al. 2016:606 [ref. 34923], Love et al. 2021:32 [ref. 39279], Page et al. 2023:51 [ref. 40505]). Current status: Valid as Anarchias galapagensis (Seale 1940). Muraenidae: Uropterygiinae. Distribution: Eastern Pacific: Pacific coast of Baja California Sur and southern Gulf of California (Mexico) south to Colombia, including Revillagigedo Islands (Mexico) and Galápagos Islands (Ecuador). Habitat: marine.
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© 2001 California Academy of Sciences

CAS 35253 Uropterygius goslinei McCosker [J. E.] & Randall [J. E.], 1977 Holotype  [Collection database record]
goslinei, Uropterygius McCosker [J. E.] & Randall [J. E.] 1977:167, Figs. 3c, 5 [Proceedings of the California Academy of Sciences (Series 4) v. 41 (no. 3); ref. 7267] Kayangel Atoll, Palau Islands, western Pacific. Holotype: CAS 35253. Paratypes: AMS I.17600-001 (1); ANSP 117434 (1), 124812 (1); BPBM 19927 (1); CAS 13972 (1), 29103 (1); USNM 215282 (1). Type catalog: Böhlke 1984:130 [ref. 13621], Smith 1994:25 [ref. 21917], Böhlke & Smith 2002:159-160 [ref. 26380]. •Valid as Uropterygius goslinei McCosker & Randall 1977 -- (Allen & Adrim 2003:23 [ref. 26830]). •Synonym of Uropterygius fasciolatus (Regan 1909) -- (see Smith 1994:25 [ref. 21917], Böhlke & Smith 2002:160 [ref. 26380], Allen & Erdmann 2012:97 [ref. 31980], Smith 2012:36 [ref. 32155], Allen & Erdmann 2024:108 [ref. 40796]). Current status: Synonym of Uropterygius fasciolatus (Regan 1909). Muraenidae: Uropterygiinae. Habitat: marine.
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CAS 35251 Uropterygius kamar McCosker [J. E.] & Randall [J. E.], 1977 Holotype  [Collection database record]
kamar, Uropterygius McCosker [J. E.] & Randall [J. E.] 1977:164, Figs. 3b, 4 [Proceedings of the California Academy of Sciences (Series 4) v. 41 (no. 3); ref. 7267] Itsandra, Grande Comore Island, Comoro Islands, Indian Ocean, depth 25 meters. Holotype: CAS 35251. Paratypes: ANSP 124794 (3), 134217 (11); CAS 35252 (20); SAIAB [formerly RUSI] 4557 (5); BPBM 12051 (1), 9442 (2), 16471 (4), 16606 (1), 17005 (1), 17047 (1); USNM 215281 (1). Type catalog: Böhlke 1984:130 [ref. 13621], Smith 1994:25 [ref. 21917], Böhlke & Smith 2002:160 [ref. 26380]. •Valid as Uropterygius kamar McCosker & Randall 1977 -- (Castle & McCosker 1986:175 [ref. 5645], Winterbottom et al. 1989:11 [ref. 13251], Myers 1999:52 [ref. 23965], Böhlke et al. 1999:1657 [ref. 24692], Randall 1999:7 [ref. 25829], Hutchins 2001:17 [ref. 25847], Böhlke & Smith 2002:160 [ref. 26380], Allen & Adrim 2003:23 [ref. 26830], Randall 2005:52 [ref. 28239], Allen et al. 2006:259 [ref. 29005], Allen & Erdmann 2012:98 [ref. 31980], Smith 2012:36 [ref. 32155], Fricke et al. 2014:21 [ref. 33932], Smith & Böhlke 2022:76 [ref. 39681], Huang et al. 2023:274 [ref. 40092], Allen & Erdmann 2024:108 [ref. 40796]). Current status: Valid as Uropterygius kamar McCosker & Randall 1977. Muraenidae: Uropterygiinae. Distribution: Indo-West Pacific: KwaZulu-Natal (South Africa), East Africa, Seychelles, Comoros and Agalega Island east to Marshall Islands and Pitcairn Islands, south to Solomon Islands. Habitat: marine.
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© 2001 California Academy of Sciences

SU 6482 Uropterygius okinawae Jordan [D. S.] & Snyder [J. O.], 1901 Holotype  [Collection database record]
okinawae, Uropterygius Jordan [D. S.] & Snyder [J. O.] 1901:886, Fig. 22 [Proceedings of the United States National Museum v. 23 (no. 1239); ref. 2508] Okinawa Island, Ryukyu Islands, Japan. Holotype (unique): CAS-SU 6482. Type catalog: Böhlke 1953:48 [ref. 12291], Böhlke & Smith 2002:162 [ref. 26380]. •Valid as Scuticaria okinawae (Jordan & Snyder 1901) -- (Böhlke & McCosker 1997:172 [ref. 23214], Fricke 1999:56 [ref. 24106], Böhlke et al. 1999:1657 [ref. 24692], Böhlke & Randall 2000:265 [ref. 24493], Nakabo 2000:198 [ref. 25086], Nakabo 2002:198 [ref. 26001], Böhlke & Smith 2002:162 [ref. 26380], Allen & Adrim 2003:23 [ref. 26830] with genus as Scuticara, Randall 2005:50 [ref. 28239], Mundy 2005:125 [ref. 28379], Randall 2007:96 [ref. 30952], Smith 2012:35 [ref. 32155], Smith & Böhlke 2022:72 [ref. 39681]). Current status: Valid as Scuticaria okinawae (Jordan & Snyder 1901). Muraenidae: Uropterygiinae. Distribution: Indo-West Pacific: Mascarenes (Mauritius) east to Hawaiian Islands (U.S.A.) and Society Islands, north to southern Japan. Habitat: marine.
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© 2001 California Academy of Sciences